IDK if this is just me, but, in some cases, the dotted lines are not enough. I become easily lost, and have to "manually" retrace the discussion.
I'd like to suggest for Tildes to use even more colors on these lines, kinda like color-schemes do for Org Mode on Emacs.
I could go even further and suggest a major "Org-Modization" of Tildes: IMHO, Org Mode has nailed this kind of structure. I know it's a bold suggestion, but there it is! ;)